ID:RN111_HUMAN DESCRIPTION: RecName: Full=E3 ubiquitin-protein ligase Arkadia; EC=6.3.2.-; AltName: Full=RING finger protein 111; FUNCTION: Acts in the NODAL pathway of mesoderm patterning during embryonic development. Acts downstream AXIN1 as an E3 ubiquitin- protein ligase which promotes the ubiquitination of inhibitory SMADs such as SMAD7, induces their proteasomal degradation and thereby enhances the transcriptional activity of TGF-beta and BMP. Activates Smad3/Smad4-dependent transcription by triggering signal-induced SnoN degradation. PATHWAY: Protein modification; protein ubiquitination. SUBUNIT: Interacts with SMAD6, SMAD7, AXIN1, AXIN2 and SKIL isoform SNON. Part of a complex containing RNF111, AXIN1 and SMAD7. SUBCELLULAR LOCATION: Nucleus. Cytoplasm. Note=Upon TGF-beta treatment, translocates from nucleus to cytosol. TISSUE SPECIFICITY: Broadly expressed. SIMILARITY: Contains 1 RING-type zinc finger. SEQUENCE CAUTION: Sequence=AAH10369.1; Type=Erroneous initiation; Note=Translation N-terminally extended; Sequence=BAC04531.1; Type=Erroneous initiation; Note=Translation N-terminally extended; Sequence=BAD18633.1; Type=Erroneous initiation; Note=Translation N-terminally extended;
The RNAfold program from the Vienna RNA Package is used to perform the secondary structure predictions and folding calculations. The estimated folding energy is in kcal/mol. The more negative the energy, the more secondary structure the RNA is likely to have.
ModBase Predicted Comparative 3D Structure on Q6ZNA4
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Orthologous Genes in Other Species
Orthologies between human, mouse, and rat are computed by taking the best BLASTP hit, and filtering out non-syntenic hits. For more distant species reciprocal-best BLASTP hits are used. Note that the absence of an ortholog in the table below may reflect incomplete annotations in the other species rather than a true absence of the orthologous gene.
Biological Process: GO:0000209 protein polyubiquitination GO:0006281 DNA repair GO:0006974 cellular response to DNA damage stimulus GO:0007275 multicellular organism development GO:0007389 pattern specification process GO:0016567 protein ubiquitination GO:0030511 positive regulation of transforming growth factor beta receptor signaling pathway GO:0030579 ubiquitin-dependent SMAD protein catabolic process GO:0031398 positive regulation of protein ubiquitination GO:0043161 proteasome-mediated ubiquitin-dependent protein catabolic process GO:0045893 positive regulation of transcription, DNA-templated GO:0045944 positive regulation of transcription from RNA polymerase II promoter GO:0070911 global genome nucleotide-excision repair
BC060862 - Homo sapiens ring finger protein 111, mRNA (cDNA clone MGC:71797 IMAGE:30336202), complete cds. BX538130 - Homo sapiens mRNA; cDNA DKFZp686H1966 (from clone DKFZp686H1966); complete cds. LF384562 - JP 2014500723-A/192065: Polycomb-Associated Non-Coding RNAs. BX647259 - Homo sapiens mRNA; cDNA DKFZp686L1262 (from clone DKFZp686L1262). AK131304 - Homo sapiens cDNA FLJ16278 fis, clone NT2RI3001132, highly similar to Mus musculus Arkadia (Arkadia) mRNA. KJ902684 - Synthetic construct Homo sapiens clone ccsbBroadEn_12078 RNF111 gene, encodes complete protein. AK131488 - Homo sapiens cDNA FLJ16671 fis, clone THYMU3001428. AL832089 - Homo sapiens mRNA; cDNA DKFZp313E0731 (from clone DKFZp313E0731). AK095327 - Homo sapiens cDNA FLJ38008 fis, clone CTONG2012452, highly similar to Mus musculus Arkadia (Arkadia) mRNA. BC020984 - Homo sapiens ring finger protein 111, mRNA (cDNA clone IMAGE:3851753), partial cds. MA620139 - JP 2018138019-A/192065: Polycomb-Associated Non-Coding RNAs. CU687480 - Synthetic construct Homo sapiens gateway clone IMAGE:100021051 5' read RNF111 mRNA. LF377550 - JP 2014500723-A/185053: Polycomb-Associated Non-Coding RNAs. JD027416 - Sequence 8440 from Patent EP1572962. JD036001 - Sequence 17025 from Patent EP1572962. LF377545 - JP 2014500723-A/185048: Polycomb-Associated Non-Coding RNAs. JD026941 - Sequence 7965 from Patent EP1572962. JD552462 - Sequence 533486 from Patent EP1572962. JD345882 - Sequence 326906 from Patent EP1572962. JD512159 - Sequence 493183 from Patent EP1572962. MA613127 - JP 2018138019-A/185053: Polycomb-Associated Non-Coding RNAs. MA613122 - JP 2018138019-A/185048: Polycomb-Associated Non-Coding RNAs. BC010369 - Homo sapiens ring finger protein 111, mRNA (cDNA clone IMAGE:4049734), complete cds. AL157474 - Homo sapiens mRNA; cDNA DKFZp761D081 (from clone DKFZp761D081). KJ902683 - Synthetic construct Homo sapiens clone ccsbBroadEn_12077 RNF111 gene, encodes complete protein. DQ583506 - Homo sapiens piRNA piR-50618, complete sequence. LF377537 - JP 2014500723-A/185040: Polycomb-Associated Non-Coding RNAs. JD192395 - Sequence 173419 from Patent EP1572962. JD398053 - Sequence 379077 from Patent EP1572962. JD113420 - Sequence 94444 from Patent EP1572962. JD402519 - Sequence 383543 from Patent EP1572962. JD113708 - Sequence 94732 from Patent EP1572962. JD181042 - Sequence 162066 from Patent EP1572962. JD563299 - Sequence 544323 from Patent EP1572962. JD489575 - Sequence 470599 from Patent EP1572962. LF377536 - JP 2014500723-A/185039: Polycomb-Associated Non-Coding RNAs. JD284229 - Sequence 265253 from Patent EP1572962. JD353837 - Sequence 334861 from Patent EP1572962. LF377535 - JP 2014500723-A/185038: Polycomb-Associated Non-Coding RNAs. LF377534 - JP 2014500723-A/185037: Polycomb-Associated Non-Coding RNAs. JD052951 - Sequence 33975 from Patent EP1572962. LF377533 - JP 2014500723-A/185036: Polycomb-Associated Non-Coding RNAs. JD327793 - Sequence 308817 from Patent EP1572962. JD473382 - Sequence 454406 from Patent EP1572962. LF377532 - JP 2014500723-A/185035: Polycomb-Associated Non-Coding RNAs. JD415458 - Sequence 396482 from Patent EP1572962. LQ283612 - Sequence 69 from Patent WO2016034611. JD090509 - Sequence 71533 from Patent EP1572962. JD189556 - Sequence 170580 from Patent EP1572962. MA613114 - JP 2018138019-A/185040: Polycomb-Associated Non-Coding RNAs. MA613113 - JP 2018138019-A/185039: Polycomb-Associated Non-Coding RNAs. MA613112 - JP 2018138019-A/185038: Polycomb-Associated Non-Coding RNAs. MA613111 - JP 2018138019-A/185037: Polycomb-Associated Non-Coding RNAs. MA613110 - JP 2018138019-A/185036: Polycomb-Associated Non-Coding RNAs. MA613109 - JP 2018138019-A/185035: Polycomb-Associated Non-Coding RNAs.
Biochemical and Signaling Pathways
Reactome (by CSHL, EBI, and GO)
Protein Q6ZNA4 (Reactome details) participates in the following event(s):
R-HSA-2186771 RNF111 binds SMAD7 R-HSA-2186785 RNF111 ubiquitinates SMAD7 R-HSA-983157 Interaction of E3 with substrate and E2-Ub complex R-HSA-983147 Release of E3 from polyubiquitinated substrate R-HSA-2186741 SMAD2/3 activation induces binding of RNF111/SMURF2 to SKI/SKIL R-HSA-2186747 Ubiquitination of SKI/SKIL by RNF111/SMURF2 R-HSA-6790487 RNF111 ubiquitinates SUMOylated XPC R-HSA-983140 Transfer of Ub from E2 to substrate and release of E2 R-HSA-2173796 SMAD2/SMAD3:SMAD4 heterotrimer regulates transcription R-HSA-983168 Antigen processing: Ubiquitination & Proteasome degradation R-HSA-2173795 Downregulation of SMAD2/3:SMAD4 transcriptional activity R-HSA-2173793 Transcriptional activity of SMAD2/SMAD3:SMAD4 heterotrimer R-HSA-5696395 Formation of Incision Complex in GG-NER R-HSA-983169 Class I MHC mediated antigen processing & presentation R-HSA-170834 Signaling by TGF-beta Receptor Complex R-HSA-212436 Generic Transcription Pathway R-HSA-5696399 Global Genome Nucleotide Excision Repair (GG-NER) R-HSA-1280218 Adaptive Immune System R-HSA-9006936 Signaling by TGF-beta family members R-HSA-73857 RNA Polymerase II Transcription R-HSA-5696398 Nucleotide Excision Repair R-HSA-168256 Immune System R-HSA-162582 Signal Transduction R-HSA-74160 Gene expression (Transcription) R-HSA-73894 DNA Repair